Ribulose 1,5-bisphosphateribulose-1 5-bisphosphate carboxylase, ribulose 1-5 bisphosphate
Ribulose 1,5-bisphosphate RuBP is an organic substance that is involved in photosynthesis It is a colourless anion, a double phosphate ester of the ketopentose ketone-containing sugar with five carbon atoms called ribulose Salts of RuBP can be isolated, but its crucial biological function happens in solution To simplify the presentation, the image in the above table depicts the acid form of this anion
Role in photosynthesisThe Calvin cycle showing the role of ribulose-1,5-bisphosphate
The enzyme ribulose bisphosphate carboxylase oxygenase RuBisCO catalyzes the reaction between RuBP and carbon dioxide The product is the highly unstable six-carbon intermediate known as 3-keto-2-carboxyarabinitol 1,5-bisphosphate This six-carbon intermediate decays virtually instantaneously into two molecules of 3-phosphoglycerate 3-PGA see figure RuBisCO also catalyzes RuBP with oxygen O
2 in a process called photorespiration, a process that is more prevalent at high temperatures During photorespiration RuBP combines with O
2 to become 3-PGA + phosphoglycolic acid In the Calvin cycle, RuBP is a product of the phosphorylation of ribulose-5-phosphate by ATP
- ^ The topic is discussed in all biochemistry textbooks, this one is representative: Nelson, D L; Cox, M M "Lehninger, Principles of Biochemistry" 3rd Ed Worth Publishing: New York, 2000 ISBN 1-57259-153-6
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