Omasumomasum, omasum of a cow
The omasum, also known as the bible, the fardel, the manyplies and the psalterium, is the third compartment of the stomach in ruminants The omasum comes after the rumen and reticulum and before the abomasum Different ruminants have different omasum structures and function based on the food that they eat and how they developed through evolution
- 1 Anatomy
- 2 Function
- 3 Species Differences
- 4 Uses in Food
- 5 See also
- 6 References
AnatomyThe 'leaflets' of the omasum
The omasum can be found on the right side of the cranial portion of the rumen The omasum receives food from the reticulum through the reticulo-omasal orifice and provides food to the abomasum through the omaso-abomasal orifice The omasum is spherical to crescent shape and has multiple leaflets similar to that of a book called omasal laminae The omasal laminae are made of thin muscular layers covered with a nonglandular mucous membrane The omasal laminae come from the sides of the large curvature and project towards the inside of the omasum, extending from the reticulo-omasal orifice to the omaso-abomasal orifice The laminae greatly increase the surface area of the omasum The laminae are covered in omasal papillae that are claw like in some ruminants or blunt cones in others These papillae further increase the surface area but they also provide increased friction against the food particles
The function of the omasum is not completely understood During the second contraction phase of the reticulum, the reticule-omasal sphincter opens for a few seconds allowing a small volume of finely dispersed and well-fermented ingesta enters the omasum
The omasum has two physiological compartments: omasal canal that transfers food from the reticulum to the omasum, and the inter-laminate recesses between the mucosal laminae which provide the area for absorption The omasum is where food particles that are small enough get transferred into the abomasum for enzymatic digestion In ruminants with a more sophisticated omasum, the large surface area allows it to play a key role in the absorption of water, electrolytes, volatile fatty acids, minerals, and the fermentation of food
Young ruminants that are still drinking milk have an esophageal groove that allows milk to bypass the rumen and go straight from the esophagus to the omasum
An early version of the omasum is seen in early ruminants like duikers and muntjacs, where it is a little more than a strainer sieve which prevents un-chewed foods from entering the abomasum
The smallest omasum belongs to ruminants that consume high quality diets like the moose and roe deer, while the largest belongs to those who are un-selective grass and roughage eaters like cattle and sheep
The omasum is not only bigger in grass and roughage eaters but there is greater differentiation in the book like structure; seen as an increase in the number of laminae
Uses in Food
See TripeSliced beef omasum is one of the Chinese dim sum known as ngau pak yip Chinese: 牛百頁 / 牛柏葉 Lap nuea dip is a northern Thai raw beef larb which includes raw beef tripe
- Methanogens in digestive tract of ruminants
- ^ a b c d The Chambers Dictionary, Ninth Edition, Chambers Harrap Publishers, 2003
- ^ a b c d e f g h Hofmann, R 1989 "Evolutionary steps of ecophysiological adaptation and diversification of ruminants: a comparative view of their digestive system" PDF Oecologia 78: 443–457 doi:101007/BF00378733
- ^ a b c d e Sjaastad, Oystein 2010 Physiology of domestic animals Hove, Knut, Sand, Olav 2nd ed Oslo: Scandinavian Veterinary Press pp 555–556 ISBN 978-82-91743-07-3 OCLC 670546738
- ^ a b c d e f Yamamoto, Y & Kitamura, N 1994 "Morphological study of the surface structure of the omasal laminae in cattle, sheep and goats" Anatomy Histology Embryology 23: 166–167 ISSN 0340-2096
- ^ a b c d Grünberg, Walter & Constable, Peter D 2009 "Function and Dysfunction of the Ruminant Forestomach" Food Animal Practice 5 ed pp 12–19 doi:101016/b978-141603591-610006-5
- ^ Braun, Ueli 2009 "Ultrasonography of the Gastrointestinal Tract in Cattle" Veterinary Clinics of North America: Food Animal Practice 25 3: 567–590 doi:101016/jcvfa200907004
- ^ Stumpff, Friederike; Georgi, Maria-Ifigenia; Mundhenk, Lars; Rabbani, Imtiaz; Fromm, Michael; Martens, Holger & Günzel, Dorothee 2011-09-01 "Sheep rumen and omasum primary cultures and source epithelia: barrier function aligns with expression of tight junction proteins" Journal of Experimental Biology 214 17: 2871–2882 doi:101242/jeb055582 ISSN 0022-0949 PMID 21832130
- ^ Green, E & Baker, C 1996 "The surface morphology of the omasum of the African goat" Journal of the South African Veterinary Association 67: 117–122 ISSN 0038-2809 PMID 9120853
- ^ a b "Animal Structure & Function" sciwaikatoacnz Retrieved 2017-11-18
- ^ "Rumen Physiology and Rumination" wwwvivocolostateedu Retrieved 2017-11-18
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