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List of animals that have been cloned

list of animals that have been clonedvd, list of animals that have been cloned animals
This is a list of animals that have been cloned The list is subheaded by animals of different types The cloned animal are included in list when citing science sources


  • 1 Camel
  • 2 Carp
  • 3 Cat
  • 4 Cattle
  • 5 Coyote
  • 6 Deer
  • 7 Dog
  • 8 Frog tadpole
  • 9 Fruit flies
  • 10 Gaur
  • 11 Goat
  • 12 Horse
  • 13 House mouse
  • 14 Monkey
    • 141 Rhesus macaque
    • 142 Crab-eating macaque
  • 15 Mouflon
  • 16 Mule
  • 17 Pig
  • 18 Pyrenean ibex
  • 19 Rabbit
  • 20 Brown rat
  • 21 Sheep
  • 22 Arctic wolf
  • 23 References
  • 24 External links


Injaz, a cloned female dromedary camel, was born in 2009 at the Camel Reproduction Center in Dubai, United Arab Emirates after an "uncomplicated" gestation of 378 days


Embryologist Tong Dizhou unsuccessfully inserted the DNA from a male Asian carp into the egg of a female Asian carp to create the first fish clone in 1963 In 1973 Dizhou inserted Asian carp DNA into a European crucian carp to create the first interspecies of this clone


  • In 2001, scientists at Texas A&M University created the first cloned cat, CC CopyCat Even though CC is an exact copy of her host, they have different personalities; ie CC is shy and timid, her host on the other hand is playful and curious
  • In 2004, the first commercially cloned cat, Little Nicky, was created by Genetic Savings & Clone


  • Gene, the first cloned calf in the world was born in 1997 at the American Breeders Service facilities in Deforest, Wisconsin, United States Later it was transferred and kept at the Minnesota Zoo Education Center Three more cloned calves were born in 1998
  • A Holstein heifer named Daisy was cloned by Dr Xiangzhong Jerry Yang using ear skin cells from a high-merit cow named Aspen at the University of Connecticut in 1999, followed by three additional clones, Amy, Betty, and Cathy in 1999
  • Second Chance, a Brahman bull was cloned from Chance, a beloved celebrity bull Second Chance was born in August, 1999 at Texas A&M University
  • Texas A&M University cloned a Black Angus bull named 86 Squared in 2000, after cells from his donor, Bull 86, had been frozen for 15 years Both bulls exhibit a natural resistance to brucellosis, tuberculosis and other diseases which can be transferred in meat
  • In 2001 researchers at Advanced Cell Technology in Worcester, Massachusetts, United States, reported that 24 successfully cloned Holsteins had been monitored from birth to the age of four All maintained healthy stats comparable to control cattle, and reached reproductive maturity at the proper stage Two of these cloned cattle successfully mated, each producing a healthy calf
  • A purebred Hereford calf clone named Chloe was born in 2001 at Kansas State University's purebred research unit This was Kansas State's first cloned calf
  • Millie and Emma were two female Jersey cows cloned at the University of Tennessee in 2001 They were the first calves to be produced using standard cell-culturing techniques
  • In 2001, Brazil cloned their first heifer, Vitória
  • Pampa, a Jersey calf, was the first animal cloned in Argentina by the company Bio Sidus in 2002
  • A Javan banteng calf was successfully cloned from frozen cells using a cow as a surrogate, delivered via c-section April 1, 2003 then hand raised at the San Diego Wild Animal Parks Infant Isolation Unit It died due to an injury when it was less than seven years old, about half the normal life of a banteng which is an endangered species
  • The world's first water buffalo was cloned in Guangxi, China by the Guangxi University in 2005 according to one reference
  • An Anatolian Grey bull Efe was cloned in Turkey in 2009 and four female calves from the same breed Ece, Ecem, Nilufer, Kiraz in 2010 by the Scientific and Technological Research Council of Turkey TÜBİTAK
  • Samrupa, the world's first Murrah buffalo a type of water buffalo calf cloned using a simple "Hand guided cloning technique" was born in 2009 at National Dairy Research Institute NDRI, Karnal, India, but died due to a lung infection five days after she was born Garima-I, a buffalo calf cloned using an "Advanced Hand guided Cloning Technique" was born in 2009 at the NDRI Two years later in 2011, she died of a heart failure Garima-II, another cloned calf was born in 2010 This buffalo was inseminated with frozen-thawed semen of a progeny tested bull and gave birth to a female calf, Mahima in 2013 A cloned male buffalo calf Shresth was born in 2010 at the NDRI
  • In May 2010, Got became the first cloned Spanish Fighting Bull, cloned by Spanish scientists
  • In February 2011, Brazil cloned a brahman
  • A Boran cattle bull was cloned at the International Livestock Research Institute in Nairobi
  • In 2015 the Chinese company BoyaLife announced that in partnership with the Korean company Sooam Biotech, they were planning to build a factory in Tianjin, China to produce 100,000 cloned cattle per year, starting in 2016 to supply China's growing market for quality beef
  • In January 2016 the scientist at the Central Institute for Research on Buffaloes in Hisar, India announced that they had cloned a buffalo offspring "Cirb Gaurav" using cells of the ventral side of the tail of superior buffalo
  • In July 2016 scientists at the National University Toribio Rodríguez de Mendoza in Chachapoyas, Peru cloned a Jersey cattle by handmade cloning method using cells of an ear of a cow The first Peruvian clon was called "Alma CL-01"


Sooam Biotech, Korea cloned eight coyotes in 2011 using domestic dogs as surrogate mothers


  • Dewey was born in 2003 at Texas A&M University


  • Snuppy, an Afghan hound puppy, was the first dog to be cloned, in 2005 in South Korea
  • Sooam Biotech, South Korea, was reported in 2015 to have cloned 700 dogs to date for their owners They were also reported to charge $100,000 for each cloned puppy One puppy was cloned from the cells of a dog that had died 12 days before
  • Sinogene, a Beijing, China-based biotechnology company, was reported in December 2017 to have cloned Apple, a gene-edited dog, with the clone being named "Longlong"

Frog tadpole

In 1958, John Gurdon, then at Oxford University, explained that he had successfully cloned a frog named Damian He did this by using intact nuclei from somatic cells from a Xenopus tadpole This was an important extension of work of Briggs and King in 1952 on transplanting nuclei from embryonic blastula cells

Fruit flies

Five genetically identical fruit flies were produced at the lab of Dr Vett Lloyd at Dalhousie University in Halifax, Nova Scotia, Canada in 2005


A species of wild cattle, the first endangered species to be cloned In 2001 at the Trans Ova Genetics in Sioux Center, Iowa, United States, a cloned gaur was born from a surrogate domestic cow mother However, the calf died within 48 hours


  • Downen TX 63 684 nicknamed Megan was cloned from a top producing Boer goat born in 2001 in Plainwell, Michigan
  • The China first cloned goat from adult ear skin,was born at Yangling,Northwest A&F University
  • The Middle East's first and the world's fifth cloned goat, Hanna, was born at the Royan Institute in Isfahan, Iran in 2009 The cloned goat was developed in the surrogate uterus of a Bakhtiari goat Iranian researchers were reported in 2009 to be planning to use cloned goats to eventually manufacture new medications such as antibodies and medicines for stroke victims
  • The world's first pashmina goat clone was produced at Centre of Animal Biotechnology at Sher-e-Kashmir University of Agricultural Sciences and Technology of Kashmir SKUAST, Indian Administered Kashmir It was named Noori, an Arabic word referring to light Funded by World Bank, this clone was a joint project of SKUAST and the Karnal-based National Dairy Research Institute


  • In 2003, the world's first cloned horse, Prometea, was born
  • In 2006, Scamper, an extremely successful barrel racing horse, a gelding, was cloned The resulting stallion, Clayton, became the first cloned horse to stand at stud in the US
  • In 2007, a renown show jumper and Thoroughbred, Gem Twist, was cloned by Frank Chapot and his family In September 2008, Gemini was born and several other clones followed, leading to the development of a breeding line from Gem Twist
  • In 2010, the first lived equine cloned of a Criollo horse was born in Argentina, and was the first horse clone produced in Latin America In the same year a cloned polo horse was sold for $800,000 - the highest known price ever paid for a polo horse
  • In 2013, the world-famous polo star Adolfo Cambiaso helped his high-handicap team La Dolfina win the Argentine National Open, scoring nine goals in the 16-11 match Two of those he scored atop a horse named Show Me—a clone, and the first to ride onto the Argentine pitch

House mouse

  • In 1986, the first mouse was cloned in the Soviet Union from an embryo cell
  • The first mouse from adult cells, Cumulina, was born in 1997 at the University of Hawai'i at Mānoa in the laboratory of Ryuzo Yanagimachi using the Honolulu technique
  • In 2008 Japanese scientists created a cloned mouse from a dead mouse that had been frozen for 16 years This was the first time a mammal had been cloned from frozen cells


Further information: wikinews:Healthy cloned monkeys born in Shanghai

Rhesus macaque

  • Tetra female, 1999 – embryo splitting artificial twinning
  • Unnamed cloned embryos 2007 – transfer of DNA from adult cells

Crab-eating macaque

  • Zhong Zhong and Hua Hua female crab-eating macaques, 2017 – first successful cloning of a primates using somatic cell nuclear transfer, the same method as Dolly, with the birth of two live female clones Conducted in China in 2017 but reported in January 2018


  • A European mouflon lamb was the first cloned endangered species to live past infancy Cloned 2001
  • A cloned baby mouflon was born to a domestic sheep in a successful interspecies cloning of an endangered species in Iran in 2015


  • Idaho Gem male, 2003 was ranked third in the world among racing mules
  • Utah Pioneer male, 2003
  • Idaho Star male, 2003


  • 5 Scottish PPL piglets Jose, Josúe, Juan, Amber and Jose 2000
  • Xena female, Meishan pig, 2000-2010
  • BGI, China was reported in 2014 to be producing 500 cloned pigs a year, with a success rate of 70-80%, to test new medicines

Pyrenean ibex

  • A cloned Pyrenean ibex was born on July 30, 2003, but died several minutes later due to physical defects in the lungs This was the first, and so far only, extinct animal to be cloned


  • In France 2003

Brown rat

  • Ralph male, 2003


  • Possibly the first cloned mammal was a sheep by Steen Willadsen in 1984 However, the cloning was done from early embryonic cells, while the sheep Dolly in 1996 was cloned from an adult cell
  • Megan and Morag were sheep cloned from differentiated embryonic cells in 1995
  • Dolly 1996–2003, first cloned mammal from adult somatic cells She had six lambs
  • Royana 2006–2010 cloned at the Royan Research Institute in Isfahan, Iran
  • Oyalı and Zarife were cloned in 2007 at Istanbul University in Istanbul, Turkey

Arctic wolf

  • The arctic wolf was cloned by South Korean scientists, including the controversial scientist Hwang Woo-Suk in 2005 The two female cloned wolves were housed in a zoo in South Korea for public view The wolves were called Snuwolf and Snuwolffy, which were names taken from Seoul National University Snuwolf died in 2009 from an infection


  1. ^ "Scientist: First cloned camel born in Dubai" The Associated Press April 14, 2009 Retrieved April 15, 2009 
  2. ^ "World's 1st cloned camel born in Dubai" Kuwait News Agency April 14, 2009 Archived from the original on April 20, 2009 Retrieved April 15, 2009 
  3. ^ Charles C Mann January 2003 "The First Cloning Superpower" Wired Retrieved 2007-06-03 
  4. ^ David Braun February 14, 2002 "Scientists Successfully Clone Cat" National Geographic Retrieved 2007-06-03 
  5. ^ "Archived copy" Archived from the original on 2014-08-08 Retrieved 2014-08-02 
  6. ^ "Pet Cat Cloned for Christmas" BBC December 23, 2004 Retrieved 2007-06-03 
  7. ^ "Calf Cloned From Bovine Cell Line" Science 277 5328: 903b–903 15 August 1997 doi:101126/science2775328903b 
  8. ^ Cibelli, J B; Stice, S L; Golueke, P J; Kane, J J 1998 "Cloned transgenic calves produced from nonquiescent fetal fibroblasts" Science 280: 1256–8 doi:101126/science28053671256 PMID 9596577 
  9. ^ "Researchers Show Clone from Aged Cow Can Produce Normal Calf" University of Connecticut web archive June 11, 2001 Retrieved 2008-05-12 
  10. ^ "Cloning gives second chance for bull" BBC News September 3, 1999 
  11. ^ Thursday, December 28, 2000 2000-12-28 "additional text" Farmanddairycom Retrieved 2012-12-11 CS1 maint: Multiple names: authors list link
  12. ^ "additional text" BBC News 2000-12-19 Retrieved 2012-12-11 
  13. ^ Whitfield, John 23 November 2001 Cloned cows in the pink Nature, retrieved 5 February 2014
  14. ^ a b Lanza, R, P, Cibelli, Jose, B, Faber, D, Sweeney, R, W, Henderson, B, Nevala, W, West, M, D, Wettstein, P, J 2011 "Cloned cattle can be healthy and normal" doi:101126/science1063440 CS1 maint: Uses authors parameter link
  15. ^ Pat Melgares May 21, 2001 "K-State's First Cloned Calf May Provide More Clues For Improving the Technology" Kansas State University Archived from the original on February 19, 2014 Retrieved 2012-12-11 
  16. ^ "Embrapa wants to fecundate Vitória, the cloned heifer" Archived from the original on 20 December 2016 
  17. ^ 6 August 002 Pampa Was Born Argentina Xplora, retrieved 29 January 2014
  18. ^ "Endangered animal clone produced" BBC News 9 April 2003 Retrieved 28 December 2015 
  19. ^ Waters, Rob 9 June 2010 "Animal Cloning: The Next Phase" Bloomberg Business Retrieved 28 December 2015 
  20. ^ "First cloned buffalo born" China View 21 March 2005 Retrieved 27 February 2015 
  21. ^ Tübitak Mam Gmbe - FK "turkhaygengovtr" turkhaygengovtr Retrieved 2012-12-11 
  22. ^ "India clones world's first buffalo" The Times Of India 
  23. ^ "Archived copy" PDF Archived from the original PDF on 2011-07-21 Retrieved 2010-05-18 
  24. ^ "First cloned buffalo dies of heart problem: NDRI scientists" The Times Of India 
  25. ^ http://wwwndriresin/ndri/Documents/cdd6c156-1297-44f5-b2b0-ada36a9f39e8pdf
  26. ^ http://wwwndriresin/ndri/Documents/779de678-8b90-46b9-b993-883e5fa9687apdf
  27. ^ "Spain clones first fighting bull" BBC 19 May 2010 
  28. ^ "First Brazilian Brahman clone born February 2011" INT'L BRAHMAN NEWS March 7, 2011 
  29. ^ Yu, M; et al 2016 "Cloning of the African indigenous cattle breed Kenyan Boran" Anim Genet doi:101111/age12441 
  30. ^ Clover, Charles; Cookson, Clive 24 November 2015 "China raises steaks in cloning research" FinancialTimes p 10 
  31. ^ DNA India - CIRB becomes India's second centre to produce a cloned buffalo, 2016
  32. ^ Diario Gestión - The first clon made in Peru was born, Alma the veal, 2016
  33. ^ Baer, Drake 8 September 2015 "This Korean lab has nearly perfected dog cloning, and that's just the start" Business Insider UK Retrieved 28 December 2015 
  34. ^ "White-tailed deer joins the clone parade - Health - Cloning | NBC News" MSNBC 2003-12-22 Retrieved 2012-12-11 
  35. ^ Mott, Maryann 3 August 2005 "Dog Cloned by South Korean Scientists" National Geographic News Retrieved 28 December 2015 
  36. ^ a b "British couple celebrate after birth of first cloned puppy of its kind" The Guardian 26 December 2015 Retrieved 27 December 2015 
  37. ^ Wang, Serenitie; Rivers, Matt; Wang, Shunhe 27 December 2017 "Chinese firm clones gene-edited dog" CNN Retrieved 29 December 2017 
  38. ^ "China clones first gene-edited dog, sentencing him to possible early death" Fox News 28 December 2017 Retrieved 29 December 2017 
  39. ^ Gurdon JB; Elsdale TR; Fischberg M 1958-07-05 "Sexually mature individuals of Xenopus laevis from the transplantation of single somatic nuclei" Nature 182 4627: 64–5 doi:101038/182064a0 PMID 13566187 
  40. ^ Robert Briggs & Thomas J King May 1952 "Transplantation of Living Nuclei From Blastula Cells into Enucleated Frogs' Eggs" Proc Natl Acad Sci U S A 38 5: 455–463 doi:101073/pnas385455 PMC 1063586  PMID 16589125 
  41. ^ "Cloning of flies is latest buzz" BBC News 4 November 2004 Retrieved 22 January 2013 
  42. ^ Advanced Cell Technology, Inc 1 December 2001 "Press Release - First cloned endangered animal was born at 7:30 PM on Monday, 8 January 2001" Archived from the original on 2008-05-31 Retrieved 2006-09-18 
  43. ^ Keith Smith - BoerGoatscom 2001-03-29 "Megan" Boergoatscom Retrieved 2012-12-11 
  44. ^ http://wwwnsfcgovcn/nsfc/cen/demonstratio/d0802htm  Missing or empty |title= help
  45. ^ "Iranian Scientists Clone Goat" CBS News Associated Press 16 April 2009 Retrieved 27 December 2015 
  46. ^ "Noori is world's first pashmina goat clone" Hindustan Times 2012-03-16 Archived from the original on 2013-01-26 Retrieved 2012-12-11 
  47. ^ Shaoni Bhattacharya August 6, 2003 "World's First Cloned Horse is Born" Retrieved 2012-05-30 
  48. ^ "Brown, Liz "Scamper Clone Offered for Commercial Breeding" ''The Horse'', online edition, November 15, 2008" Thehorsecom 2008-11-15 Retrieved 2012-12-11 
  49. ^ "Clone of top jumper Gem Twist born" horsetalkconz September 17, 2008 
  50. ^ Andrés Gambini Javier Jarazo Ramiro Olivera Daniel F Salamone 2012 "Equine Cloning: In Vitro and In Vivo Development of Aggregated Embryos" Biol Reprod doi:101095/biolreprod112098855 CS1 maint: Uses authors parameter link
  51. ^ Cohen, Haley 31 July 2015 "How Champion-Pony Clones Have Transformed the Game of Polo" VFNews Vanity Fair Retrieved 27 December 2015 
  52. ^ Alexander, Harriet 8 December 2014 "Argentina's polo star Adolfo Cambiaso - the greatest sportsman you've never heard of" The Telegraph Retrieved 27 December 2015 
  53. ^ Ryan Bell "Game of Clones" Outside Online 
  54. ^ Six cloned horses help rider win prestigious polo match - Jon Cohen, Science Magazine, 13 December 2016
  55. ^ Chaĭlakhian LM, Veprintsev BN, Sviridova TA 1987 "Electrostimulated cell fusion in cell engineering" Biofizika 32 5: viii–xi 
  56. ^ Nowak, Rachel 3 November 2008 Cloning 'resurrects' long-dead mice The New Scientist, Retrieved 13 April 2014
  57. ^ "Scientists 'clone' monkey, BBC News" 14 January 2000 
  58. ^ "Cloned monkey stem cells produced" Nature News 22 November 2007 
  59. ^ Liu, Zhen; et al 24 January 2018 "Cloning of Macaque Monkeys by Somatic Cell Nuclear Transfer" Cell doi:101016/jcell201801020 Retrieved 24 January 2018 CS1 maint: Explicit use of et al link
  60. ^ Briggs, Helen 24 January 2018 "First monkey clones created in Chinese laboratory" BBC News Retrieved 24 January 2018 
  61. ^ Associated Press 24 January 2018 "Scientists Successfully Clone Monkeys; Are Humans Up Next" The New York Times Retrieved 24 January 2018 
  62. ^ "Chinese Scientists Clone Monkeys Using Method That Created Dolly The Sheep" NPR January 24, 2018 
  63. ^ "Scientists Clone First Endangered Species: a Wild Sheep" Newsnationalgeographiccom 2010-10-28 Retrieved 2012-12-11 
  64. ^ Deghan, Saeed Kamali 5 August 2015 "Scientists in Iran clone endangered mouflon – born to domestic sheep" The Guardian Retrieved 27 December 2015 
  65. ^ Black, Richard May 29, 2003 "Cloning first for horse family" BBC 
  66. ^ McManus, Phil; Albrecht, Glenn; Graham, Raewyn 31 August 2012 The Global Horseracing Industry: Social, Economic, Environmental and Ethical Perspectives Routledge p 178 ISBN 978-0415677318 
  67. ^ "Pigs cloned for organs down at the 'pharm'" The Guardian London March 14, 2000 
  68. ^ "Research progress: Pig cloning for organs" CNN January 3, 2002 Archived from the original on October 5, 2012 
  69. ^ Shukman, David 14 January 2014 China cloning on an 'industrial scale' BBC News Science and Environment, Retrieved 14 January 2014
  70. ^ J Folch; J Cocero; M J Chesne; P Alabart; J K Dominguez; V Congnie; Y Roche; A Fernández-Árias; A Marti; J I Sánchez; P Echegoyen; E Beckers; J F Sánchez; A Bonastre; X Vignon 2009 "First birth of an animal from an extinct subspecies Capra pyrenaica pyrenaica by cloning" Theriogenology 71 6: 1026–1034 doi:101016/jtheriogenology200811005 PMID 19167744 
  71. ^ Zimmer, Carl "Bringing Them Back To Life" Retrieved September 13, 2014 
  72. ^ Challah-Jacques, M; Chesne, P; Renard, J P 2003 "Production of Cloned Rabbits by Somatic Nuclear Transfer" Cloning and Stem Cells 5 4: 295–299 doi:101089/153623003772032808 PMID 14733748 
  73. ^ First cloned rabbit, Stice L, Steven PhD "Unlocking the Secrets of Science" PDF University of Georgia 2010 
  74. ^ "Ralph: The World's First Cloned Rat" Brighthubcom Retrieved 2012-12-11 
  75. ^ Bartlett, Zane, "Somatic Cell Nuclear Transfer in Mammals 1938-2013" Embryo Project Encyclopedia 2014-11-04 ISSN: 1940-5030 http://embryoasuedu/handle/10776/8231
  76. ^ "Dolly the Sheep" University of Edinburgh, Roslin Institure 7 April 2015 Retrieved 9 April 2015 
  77. ^ Sheikhi, Marjohn 30 September 2015 "Birth anniversary of Royana; Iran's 1st cloned sheep" Mehr News Agency Retrieved 2017-12-09 
  78. ^ "Türkiye´nin ilk kopya koyunu doğdu - BİLİM-TEK Haberleri" Haber7com 2008-06-08 Retrieved 2012-12-11 
  79. ^ "Turkey's first cloned sheep born at Istanbul University | Science | RIA Novosti" Enrianru 2007-11-22 Retrieved 2012-12-11 
  80. ^ "Türkiye´nin 2 kopya koyunu Zarife - BİLİM-TEK Haberleri" Haber7com 2008-06-08 Retrieved 2012-12-11 
  81. ^ "Not Extinct Yet : Snuwolf and Snuwolffy" 
  82. ^ "World's first cloned wolf dies" PhysOrg 1 September 2009 Retrieved 9 April 2015 

External links

  • "Gallery of Cloned Animals" Guardian Unlimited Archived from the original on 12 March 2016 
  • "UTK clone project" Knoxville, TN Archived from the original on 9 February 2012 
  • Iran's first cloned goat born in Isfahan
  • "Equine Cloning: In Vitro and In Vivo Development of Aggregated Embryos" Biol Reprod 2012 doi:101095/biolreprod112098855 

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